| Pattern formation out of equilibrium is intensively studied topic. This is because that can explain the dynamical spatio-temporal organization of biological cells, cell colonies and tissues. In this study, we present an experimental system where large scale patterns emerge from local interaction of propelled biological filaments. The system is an in vitro motility assay where grafted motor proteins dynein c propel microtubules. We observed collision behavior of microtubules, which leads to direct alignment. At large enough density of microtubules, we observed large vortices formed by microtubules moving both clockwise and anticlockwise. A simple mathematical model similar to the Vicsek model but specifically designed for this experimental situation successfully reproduces the observed patterns and offers predictions for even large time- and length-scale behaviour of this system. |
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